This is a nice simple little discussion of ant navigation while dragging food home backwards.
I actually was wondering about this as I watched a Veromessor pergandei dragging home dandelion seeds (and missing the test tube entrance due to a piece of fluff). Clearly there are still lots of questions remaining, though.
How ants walking backward find their way home
https://www.sciencem...-their-way-home
Past & Present
Veromessor pergandei, andrei, stoddardi; Novomessor cockerelli
Camponotus fragilis, Camponotus sansabeanus (inactive), vicinus, laevigatus/quercicola, CA02
Pogonomyrmex subnitidus, P. californicus (inactive)
Liometopum occidentale (inactive); Prenolepis imparis; Myrmecocystus mexicanus (inactive); Tetramorium sp. (inactive); Lasius sp.
Termites: Zootermopsis angusticollis, and a box of drywood termites that can't be seen
Isopods: (most no longer keeping) A. gestroi, granulatum, kluugi, maculatum, vulgare; C. murina; P. hoffmannseggi, P. haasi, P. ornatus; V. parvus, P. pruinosus, T. tomentosa
Spoods: (no longer keeping) Phidippus sp., other